Fish fossil remains from the Khartam Member, Khuff Formation in Central Saudi Arabia: Implications for placement of the Permo/Triassic Boundary

Recognition of the Permian/Triassic boundary in Saudi Arabia by previous researchers using a variety of methods has resulted in placement of the boundary at different stratigraphic levels. In this study, we discovered two lithological horizons containing fossil fish teeth (sharks and Actinopterygii) in different stratigraphic positions within the Khartam Member of the Khuff Formation. Thin-section petrography initially revealed the fossil fish teeth, while XRF, XRD, SEM, and QEMSCAN techniques were used to confirm the presence of biogenic apatite. Fish teeth collected from the two stratigraphic levels in the Khartam Member show distinct and different morphological features. The fossil assemblage recovered from the lower unit belongs to the classes Euselachii and Actinopterygii, while fish teeth from the upper unit belong to the shark genus Lissodus.  The stratigraphic position of the recovered Lissodus microfossils supports the previous placement of the Permo/Triassic boundary by Vaslet et al. (2005) and Crasquin-Soleau et al. (2006), who placed the boundary exactly at the stratigraphic boundary between the Lower and Upper Khartam Members. The results of this study do not corroborate the boundary placement by Eltom et al. (2016), who placed the Permo/Triassic boundary in a higher stratigraphic position within the Upper Khartam Member. Lissodus species are common from the Early Triassic to Late Cretaceous, suggesting new paleontological evidence for the placement of the Permo/Triassic boundary in Saudi Arabia.